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view on githubraw file Latest commit 8916dab9 on 2022-03-21 14:59:06 UTC
8916dab93b Oliv*0001 .. include:: ../defs.hrst
                0002 
                0003 .. _CarbonStore:
                0004 
                0005 Internal carbon store and exudation
                0006 '''''''''''''''''''''''''''''''''''
                0007 
                0008 Note: the internal carbon store is currently only implemented for fixed
                0009 plankton elemental ratios (i.e., :varlink:`DARWIN_ALLOW_NQUOTA`,
                0010 :varlink:`DARWIN_ALLOW_PQUOTA`, :varlink:`DARWIN_ALLOW_FEQUOTA`,
                0011 :varlink:`DARWIN_ALLOW_SIQUOTA` all undefined).
                0012 
                0013 With :varlink:`DARWIN_ALLOW_CSTORE` defined, nutrient availability limits
                0014 growth directly rather than via the photosynthesis rate.  The maximum
                0015 photosynthesis rate (and with it carbon uptake) becomes independent of
                0016 nutritient limitation,
                0017 
                0018 .. math::
                0019 
                0020    P^{{\mathrm{C}}{\op{m}}}_j = P_{{\mathrm{C}},j}^{\max} f^{{{\text{phy}}}}_j(T) \gamma_{\op{pCO2}}
                0021 
                0022 but the growth rate is limited by the availability of each nutrient,
                0023 
                0024 .. math::
                0025 
                0026    \mu_j = \min(P^{\op{C}}_j, \mu^{\max\op{N}}_j, \mu^{\max\op{P}}_j,
                0027            \mu^{\max\op{Fe}}_j, \mu^{\max\op{Si}}_j)
                0028 
                

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0029 Each nutrient’s availability determines a maximum achievable growth rate. Its 0030 form depends on whether variable quotas are enabled for the nutrient. 0031 For instance for phosphorus, without P quota, 0032 0033 .. math:: 0034 :label: eq_darwin_max_growth_p 0035 0036 \mu^{\max\op{P}}_j = \frac{1}{R^{\op{P}:\op{C}}_j} V^{{\mathrm{P}}\max}_j 0037 \gamma^{\op{P}}_j f^{\op{up}}_j(T) ( 1 + r^{\op{resp}}_j ) 0038 0039 and with P quota, 0040 0041 .. math:: 0042 :label: eq_darwin_max_growth_p_noquota 0043 0044 \mu^{\max\op{P}}_j = \frac{1}{R^{\op{P}:\op{C}}_j} u^{\mathrm{P}}_j 0045 ( 1 + r^{\op{resp}}_j ) 0046 0047 where :math:`u^{\mathrm{P}}_j` is the carbon-specific uptake rate for phosphorus, 0048 0049 .. math:: 0050 0051 u^{\mathrm{P}}_j = V^{{\mathrm{P}}\max}_j \frac{\mathrm{PO}_4}{\mathrm{PO}_4 + k^{\op{PO4}}_j} 0052 {{\text{reg}}}^{Q{\mathrm{P}}}_j \cdot f^{{\text{up}}}_j(T) 0053 0054 and the respiration rate is 0055 0056 .. math:: 0057 0058 r^{\op{resp}}_j = r^{\op{resp}\max}_j f^{\op{remin}}(T) \frac{c_j - c_j^{\min}}{c_j} 0059 0060 Other elements work analogously, except that the maximum nitrogen uptake rate 0061 for the case without nitrogen quotas is more complex, 0062 0063 .. math:: 0064 0065 \mu^{\max\op{N}}_j = \frac{1}{R^{\op{P}:\op{C}}_j} u^{{\mathrm{N}}\max}_j 0066 ( 1 + r^{\op{resp}}_j ) 0067 0068 where 0069 0070 .. math:: 0071 0072 u^{{\mathrm{N}}\max}_j = \max( u^{{\mathrm{NO3}}\max}_j, 0073 u^{{\mathrm{NO2}}\max}_j, u^{{\mathrm{NH4}}\max}_j ) 0074 0075 with 0076 0077 .. math:: 0078 0079 u^{{\mathrm{N..}}\max}_j = V^{{\mathrm{N..}}\max}_j \gamma^{\op{N..}}_j 0080 f^{\op{up}}_j(T) 0081 0082 Part of the excess carbon is exuded to become DOC, 0083 0084 .. math:: 0085 0086 E^{\op{C}}_j = (P^{\op{C}}_j - \mu_j) f^{\op{exude}}_j c_j 0087 0088 and part stored as fat, 0089 0090 .. math:: 0091 0092 U^{\op{fat}}_j = (P^{\op{C}}_j - \mu_j) (1 - f^{\op{exude}}_j) c_j \;. 0093 0094 The Chlorophyll synthesis rate is based on the growth rate (as before). 0095 0096 .. csv-table:: Exudation parameters 0097 :delim: & 0098 :widths: 13,20,17,15,15,20 0099 :class: longtable 0100 :header: Trait, Param, Symbol, Default, Units, Description 0101 0102 :varlink:`FracExudeC` & :varlink:`a_FracExudeC` & :math:`f^{\op{exude}}_j` & 0.3 & & fraction of excess carbon exuded